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- B.S., Engineering (Aeronautics), University of Tokyo, Tokyo, Japan, (1988-1992)
- M.S., Biological Science, Nara Institute of Science and Technology, Nara, Japan, (1995-1997)
- Ph.D., Medicine (Neuroscience), University of Tokyo Graduate School of Medicine, Tokyo, Japan, (1997-2001)
- Post Doc, Pharmacology, Vanderbilt University, Nashville, TN, (2002-2008)
- Post Doc, Neurobiology, Yale University School of Medicine, New Haven, CT, (2005-2008)
